Cross-appointed in Dalhousie’s Department of Biochemistry and Molecular Biology and its Department of Philosophy, successful candidates will investigate applications of the theory of evolution (broadly construed) and develop new understandings of mutualisms, multi-species symbioses and communities. We are specifically interested in extensions of Darwinian approaches that embrace persistence, niche construction, Hull’s replicator/interactor framework, evolutionary individuality, process ontology, lineage concepts and cooperation, from holobionts to biogeochemical cycles and the biosphere. This work will be supported by funds made available through the Gordon and Betty Moore Foundation’s Symbiosis in Aquatic Systems Initiative. Supervision will be by W. Ford Doolittle (email@example.com), who is PI on this project, and fellows will interact with an existing team comprising two philosophers, a population biologist, and a computational modeler. One goal would be a deeper evolutionary understanding of the rapidly accumulating databases resulting from marine microbial metagenomic research, and familiarity with methods for analyzing and modeling such data would be an asset, but not a requirement.